"I do not care about Polanski, but I hope that if my daughter is ever raped it is for an older and truly talented man with a great sense of staging," Mr. Seder's tweet said.
Eight years later, Mr. Cernovich, an activist who promoted the #Pizzagate conspiracy theory, helped draw attention to the tweet and began harassing the MSNBC media relations department and other journalists about Mr. Seder.
Mr. Cernovich did not immediately respond to a request for comment.
In his podcast, Seder accused Mr. Cernovich of trying to silence his frequent criticisms of President Trump and Alabama Senate candidate Roy Moore.
After MSNBC fired Mr. Seder, thousands of people signed a petition in protest, arguing that Mr. Cernovich had acted in bad faith, taking Mr. Seder's tweet, which was meant to be a satire, out of context.
Chris Hayes, one of the network's star hosts, went public with his anger over Mr. Seder's treatment of MSNBC, though he did not mention Mr. Seder by name.
"I reiterate my long-standing position that people should not be fired by a tweet, * particularly * one that obviously reads with manifest bad faith," Mr. Hayes tweeted on Monday.
On Thursday, MSNBC regained its position.
"We made our initial decision for the right reasons, because we do not consider rape to be a fun subject to be joked about," Mr. Griffin said in his statement. "But we've heard the comments, and we understand that the point Sam was trying to make in that tweet was actually in line with our values, even though the language was not, Sam will be welcome in our air in the future."
Mr. Seder also issued a statement saying: "I appreciate the careful reconsideration of MSNBC and its willingness to understand the cynical motives of those who intentionally misrepresented my tweet for their own toxic and political purposes."
He explained in a telephone interview, saying about MSNBC: "I think they were wrong, and I think hopefully they learned a lesson that you have to do an evaluation on the substance." The media in general have been very reluctant to do so. , and I think it has become much more important in this era, both because of technology and because of the depravity of some elements of our society. "
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